The LGB chassis is effectively a Mallet. We have a 6 wheel fixed motor block and a flexible 4 wheel motor block to the rear (plus the pony truck - but I'll ignore those for this) Each block has its own motor. They are connected by a universal joint drive to keep the speeds the same and to allow the coupling rods to stay in place in relation to the wheels. The hole in the connecting rod on the middle drivers is elongated to allow the rear motor block to swing.
On the original, the front two axles are allowed to have a lot of sideplay( in the rigid chassis). This is controlled by the levers which allow the front axle to move one way and push the second axle the other. The advantage here is not only flexibility for cornering but also puts both the first and second axles flange against the rail to give smoother turning force (also safer - less likely to push the first axle over the rail head.) The crank pin journals have similarly excessive sideplay to allow the coupling rods to remain straight.
So, back to your question - the LGB loco allows the rear 4 drivers to move sideways and the original allows the front 4 wheels to move sideways.
Clear as mud?